Meet the Team

The Wilcove Team
(aka The Drongos)

David S. Wilcove (professor): Members of my research group use a combination of ecology, economics, and policy research to find innovative ways to protect biodiversity around the world. Recent or ongoing projects include: studies of the impact of logging and oil-palm agriculture on biodiversity in Southeast Asia; impact of the cage-bird trade on Indonesian birds; conservation of migratory animals; factors affecting the distribution of birds along elevational gradients in the Himalayas; reforestation and forest recovery in China, Costa Rica, and Peru; and the impacts of hunting on wildlife in China and Brazil. Prior to joining the Princeton faculty in 2001, I served as senior ecologist with the Environmental Defense Fund (1991-2001) and The Wilderness Society (1986-1991). I am an avid birdwatcher who cannot forgive himself for missing the Hawaiian Crow in 1996, but who did see the last wild Spix’s Macaw in Brazil in 1993.

Dan Liang with backpackDan Liang (postdoc): I am a field ecologist with broad interests in ecology, evolution, and conservation biology. My main research aims to understand how human-induced disturbances impact biodiversity. My PhD work looked at how urbanization affects stress responses and various levels of niche characteristics of passerine birds in subtropical environments in Southwest China. For my postdoc, I am working with Prof. David Wilcove to try to understand the effects of hunting or trapping on populations of birds, especially those birds migrate along the East Asian- Australasia Flyway.

Prior to Princeton, I received my PhD degree in Zoology from Sun Yat-sen University (2018), and received my master degree in Zoology (2015) and my bachelor degree in Wildlife and Nature Reserve Management (2012) from Southwest Forestry University in China. Read more about my research on my website:


Liang MaLiang Ma (postdoc): Before I became a Drongo, I focused my research on understanding the relationship between thermal variation over space and time and the physiology and life history of ectotherms. I parameterized mechanistic models with reaction norms derived from empirical studies to generate new insights about the process of thermal adaptation at both local and global scales. Now, I am exploring how climate and land-use change will impact pikas in China. Specifically, I currently plan to answer this question from the perspectives of species distribution and physiological limitation, respectively.

From 2012 to 2019, I completed my PhD and did a postdoc with Wei-guo Du at the Institute of Zoology, Chinese Academy of Sciences. I received my BS from Beijing Forestry University in 2012. During 2015-2016, I did a one-year internship at the University of Washington (Seattle) with Lauren B. Buckley and Raymond B. Huey.

Tong Mu (postdoc): My research interests lie in the field of avian ecology and conservation, especially that of shorebirds migrating along the East Asian-Australasian Flyway. Combining field study, individual tracking and mathematical modeling, I plan to identify the factors affecting their selections of migration routes and stopover sites, and to see whether the understanding of shorebirds migration patterns can be extended to that of other migratory birds.

Prior to joining the Drongos, I received my B.S. from Peking University in 2014, and during my time there, I became an avid birdwatcher and also became aware of the conservation issues pertaining to birds and other organisms.


Yiwen ZengYiwen Zeng (postdoc): I am an applied ecologist and geospatial modeller. My research focuses on tropical conservation and sustainability, with the overarching goal of applying interdisciplinary solutions to issues surrounding environmental sustainability and its intersection with human development, climate change and biodiversity conservation.




Chris CrawfordChristopher Crawford (graduate student): Some of my research interests include ecological restoration (particularly along rivers and streams), incentives for conservation and restoration on private lands, the effects of agricultural land use on biodiversity, and causes and consequences of land use change. Big questions include: how do we find more room for biodiversity in human-dominated landscapes, and how do we feed the world without destroying it? By exploring the biodiversity, ecosystem service, and economic trade-offs of different land use scenarios, and looking at policies and incentives to effectively guide land use, I hope to improve how we use land for conservation and for people.

Prior to graduate school, I spent 3.5 years working for Sustainable Conservation in San Francisco, on collaborative projects to 1) reduce the use of invasive plants in horticulture (PlantRight) and 2) facilitate riparian restoration in California’s Central Valley. I graduated from the University of Michigan in 2012, and I love riding my bicycle, taking pictures of clouds, and listening to Swedish music.

freda guoFreda Fengyi Guo (graduate student): I’m a conservation ecologist interested in ecological principles and applying them to conservation practices. My research interests cover a wide range of conservation-related topics including the stopover ecology of migratory birds and the conservation implications, species’ regional movement patterns and community structures in fragmented habitats, as well as impacts of China’s emerging ecotourism on conservation.

Prior to Princeton, I completed my BSc and MPhil degrees at the University of Hong Kong. For my MPhil, I studied interactive effects between climate and land-use change on species elevational range shifts, and behavioral thermal constraints on global ants’ distributions. I enjoy reading, hiking, photography, and Chinese calligraphy. I’m also a Spurs fan in case you want to talk about basketball :)

Brian LeeBrian Lee (graduate student): My research interests lie at the intersection of agricultural economics and conservation science. Specifically, I hope to apply methods from agricultural economics to answer questions related to biodiversity. Some of my previous work examined the causal impact of afforestation programs on income inequality among forest farm households in Taiwan. I have also examined the causal impact of COVID-19 on metro use in Taiwan. In the future, the research questions that I hope to answer include how conservation policies affect farmland and farm households. I am also interested in studying the impact of agricultural production on biodiversity.

I completed a M.Sc. in Agricultural Economics on a Fulbright Grant at National Taiwan University in Taipei, Taiwan. I also have a B.S. in Economics from the University of California, Davis.


bing linBing Lin (graduate student): Broadly speaking, I am interested in the areas of overlap between conservation ecology, behavioral economics, and public policy. Using insights from the behavioral sciences, I want to conduct research on the manifold ways science (and policy) can provide simple yet effective solutions to problems in ecology, biology, and the environment. From fishermen to college freshmen to you and me, we all stand to gain from our planet's well-being and suffer in its deterioration. As such, my research will hopefully tackle the alignment issue of how we can make the right thing to do the easy thing to do in our lives.

I obtained an undergraduate degree in Ecology and Evolutionary Biology from Princeton in 2016, where I conducted research assessing the relative roles of fish assemblages and sea urchins (Diadema antillarum) on Caribbean coral reef health for my Senior Thesis. In the time following, I spent a year on a Princeton in Asia fellowship teaching at a boarding school in Thailand, a year conducting behavioral research on gelada monkeys (Theropithecus gelada) in the highlands of Ethiopia, and several months surveying coral reefs in Dominica and hiking the Pacific Crest Trail in western North America. In my spare time, you can find me scuba diving, snowboarding, hiking, taking pictures of wildlife, or playing pretty much any sport.

Alex WiebeAlex Wiebe (graduate student): I am a biologist interested in global changes in biodiversity. I study how biodiversity loss is tied to global development across scales, from mechanisms of gradual species loss under local reductions in habitat area to patterns of biodiversity loss associated with trade globalization.

My research is split between field ecology and modeling and involves experimental systems in bird communities in the Brazilian Amazon. Before joining the Drongos, I graduated from Cornell University with a BS in Biology.


drongos research group zoom photo
August 26, 2020: Drongos from three continents convene for a virtual group meeting during the pandemic.

February 19, 2020: the full Drongo flock












March 2018 Drongos
March 14, 2018: Another rare sighting of all of the Drongos in the same place at the same time.













On a cold day in March 2014, all of the Drongos were together in Princeton for the first time in over a year. Todd Reichart (Chemistry Department) took this historic photo to commemorate the event.





This photo from February 2013 represents a rare occasion when all of the Drongos were together at the same time on the Princeton Campus. Usually at least one of us is in Indonesia, Malaysia, Brunei, China, India, Korea, Peru, Ecuador, or even Tennessee.






Drongo Alumni


Bethany Bradley (former postdoc): Bethany studied the impact of climate change on invasive plants for the Western USA from 2006-2006. She is now at the University of Massachusetts at Amherst.

Zuzana Burivalova (former postdoc): Zuzana focused on innovative ways to prioritize sites for conservation in Southeast Asia and New Guinea using a combination of fieldwork and data-mining. She is now at the University of Wisconsin - Madison.

Charlotte Chang (former graduate student): Charlotte studied the ecological and social dynamics of hunting in Yunnan. She is now at the University of Tennessee.

Willandia Chaves (former postdoc): Willa studied how urbanization affects the consumption of wild animals as food for people in the Brazilian Amazon from 2017-2020. She is now at Virginia Tech.

David Edwards (former postdoc): David studied patterns of biodiversity in primary once-logged and twice logged forests in Southeast Asia. He is now at the University of Sheffield.

Paul Elsen (former graduate student): Paul studied the ecology and conservation of bird communities in the Himalayas. He is now at the Wildlife Conservation Society.

Lyndon Estes (former postdoc): Lyndon modeled how climate change is likely to shift maize and wheat cultivation in South Africa, as well the implications of those shifts to biodiversity. He is now at Clark University.

Brendan Fisher (former postdoc): Brendan studied the economics of logging and oil palm agriculture in Southeast Asia. His is now at the University of Vermont.

Eyal Frank (former postdoc): Eyal studied the protection of wildlife threatened by international trade. He is now at the University of Chicago.

Xingli Giam (former graduate student): studied the conservation of freshwater biodiversity in Southeast Asia from 2009-2014. He is now at the University of Tennessee-Knoxville.

Jonathan Green (former postdoc): studied the ecological and socioeconomic issues related to the conservation of shorebirds (e.g. spoon-billed sandpiper) wintering in Southeast Asia during 2012-2014. He is now at the University of York.

Nathan Gregory (former graduate student): Nathan studied how prescribed fire and Massai pastoralism affected bird diversity in the Sayannas of East Africa. He is now at the Irvine Ranch Conservancy.

Bert Harris (former postdoc): studied the impact of the bird trade in populations of wild birds in Indonesia from 2012-2015. He is now at the Clifton Institute.

Josh Hooker (former postdoc): From 2005-2008, Josh studied the impacts of climate change on bird communities in North America. He is now at the University of Reading.

Fangyuan Hua (former postdoc): Fangyuan studied the biodiversity impacts of China's reforestation programs. She is now at Peking University.

Lian Pin Koh (former graduate student): Pin was a graduate student from 2004-2008, studying the impacts of oil-palm agriculture on biodiversity in Southeast Asia. He is now at the National University of Singapore.

Trond Larsen (former postdoc ): Trond was a WWF Fuller Postdoctoral Fellow from 2008-2010. He used dung beetles as a model system for developing conservation strategies in the Andes-Amazonia region. He is now at Conservation International.

Janice Ser Huay Lee (former postdoc): Janice studied the expansion of oil palm agriculture in Southeast Asia and its impacts on biodiversity. She is now at the Nanyang Technological University, Singapore.

Tien Ming Lee (former postdoc): Ming studied the social drivers of the wild bird trade in Indonesia. He is now at Sun Yet-sen University.

Dave Marvin (former research assistant): From 2006-2008, Dave created a novel web-mapping system to collect data on the distribution and abundance of invasive plants in the Southeast U.S. He is nowat The Nature Conservancy.

Emily Nicholson (former postdoc): Emily was a postdoc from 2006-2007. Working in collaboration with The Nature Conservancy and the Florida Natural Areas Inventory, she developed new quantitative methods for assessing progress in biodiversity conservation. She is now at Deakin University.

David Pattemore (former graduate student): David studied how the loss of native vertebrates in New Zealand has affected the pollination ecology of the native plants there; he also studied the degree to which non-native vertebrates are assuming the pollination roles of the missing natives. He is now with Plant & Food Research in New Zealand.

Rebecca Senior (former postdoc): Rebecca's main project analyzed data from the IUCN Red List to understand which conservation actions have been successful in recovering threatened species. She is now at Durham University.

Jacob Socolar (former graduate student): Jacob studied the impacts of land-use change on Amazonian birds. He is now at NCX.

Umesh Srinivasan (former postdoc): Umesh studied the ecology and conservation of Himalayan birds. He is now at the Centre for Ecological Sciences, Indian Institute of Science.

Morgan Tingley (former postdoc): explored the factors driving long-term changes in the avifauna of the Great Smokies Mountains from 2012-2014 with support from David H. Smith Conservation Research Fellowship Program. He is now at UCLA.

Tim Treuer (former graduate student): Time studied the factors affecting natural forest restoration in Costa Rica. He is now at the University of Vermont.

Will Turner (former postdoc): Will was a post-doctoral fellow from 2003-2006, working on issues pertaining to reserve design and management. He is now at Conservation International.

Charles Yackulic (former postdoc): Charles modeled the spatial and population dynamics of spotted owls and barred owls in the Pacific Northwest. He is now with the USGS Biological Resources Division.